How to Play

Dominant Species: Marine

2-4 players 90-150 min 7.9 Board Game Geek Rating

Players control aquatic animal classes (reptiles, fish, cephalopods, crustaceans) competing for dominance after an ice age ends. Through strategic action pawn placement, species adapt, migrate, and thrive across expanding ocean habitats. Victory comes from accumulating the most points by dominating elements, controlling territories, and collecting powerful cards before an asteroid arrives.

Published 2021

Dominant Species: Marine

Get Ready

Setup the game with these step-by-step instructions before diving into your turn.

1
Place the board
Put the main board in the center of the table
2
Arrange initial terrain tiles
Place five Land tiles, five Open Ocean tiles, and one each of the other five types in their assigned positions at the center of the grid
3
Create terrain draw piles
Flip remaining 21 large tiles face down, shuffle them, create three piles of 7 tiles each on the Exploration spaces, then flip the top tile of each pile face up
4
Place Hydrothermal Vent tiles
Put one Hydrothermal Vent tile (submarine side up) on the lowest Open Ocean tile and another (land side up) on the highest Land tile, then place remaining 10 Vent tiles in their designated stack
5
Set up starting elements
Take one Element disc of each of the six types and place them around the central Coral Reef tile as indicated, covering parts of all three tiles at each intersection
6
Fill element bag
Place all remaining Element discs in the black bag
7
Populate action sections
Draw 4 random Elements and place them in the Abundance section's jellyfish circles, then repeat for Adaptation, Speciation, and Exploration sections
8
Set up terrain tokens
Put all 16 Terrain tokens in the red bag, draw 3 random tokens for the Competition section's starfish spaces, then draw 5 more tokens for the Evolution section arranged left to right by type
9
Distribute Animal Boards
Give each player the Animal Board matching their chosen animal, using the side with three pre-printed elements
10
Assign action pawns
Each player takes colored wooden cylinders based on player count: 4 cylinders for 4 players, 5 for 3 players, or 7 for 2 players
11
Apply stickers
Before first game, apply matching sticker pairs to top and bottom of each white cylinder
12
Place Control tokens
Put the six Control tokens on their designated board spaces with matching icons, then place corresponding Special Pawns on top
13
Position Threshold markers
Place all six Element Threshold tokens on space "1" of the Victory Point track
14
Initialize score markers
Place each animal's VP marker on space "0" of the track with the +100 side face down
15
Prepare species cubes
Each player places their 35 colored cubes in the Gene Pool section of their Animal Board
16
Set Reseed markers
Each player takes 1 cube from their Gene Pool and places it in the left box next to their animal icon on the Food Chain table
17
Place starting species
Each player places 3 cubes from their Gene Pool onto the central Coral Reef tile
18
Draft Trait cards
Shuffle the 18 Trait cards face down, deal 3 to each player, return the rest to the box unseen, then each player selects 1 card to keep and discards the others
19
Reveal Trait cards
All players reveal their chosen Trait card and place it face up on their Animal Board
20
Prepare Evolution deck
Remove the Asteroid card, shuffle remaining 34 cards face down, remove 10 cards unseen, remove 4 more cards, shuffle Asteroid into those 4 cards, place this 5-card packet face down beside the board, then place remaining 20 Evolution cards face down on top
21
Display Evolution cards
Draw the top 5 Evolution cards and place them face up in the Evolution card spaces on the left side of the board
22
Check entry effects
Resolve any "When this card enters play" effects on revealed Evolution cards
23
Set aside Survival card
Place the Survival card to the side for now
24
Determine first player
The animal lowest on the Food Chain takes the first turn

On Your Turn

What you can do during your turn.

1
Choose one option
During your turn, you must either place a pawn to take an action OR retrieve all your pawns
2
Place a pawn
Put one of your available pawns (colored Base Pawn or white Special Pawn you control) on a single fossil action space and immediately perform the associated action
3
Retrieve pawns
Take all your controlled pawns (both Base and Special) from the Action Area and place them in front of you as available pawns for future turns
4
Base Pawn placement restrictions
Your Base Pawns may only be placed in empty action spaces that are not Special Pawn exclusive and are positioned "after" your other pawns
5
Special Pawn flexibility
Special Pawns can be placed in any empty action space OR in spaces containing an opponent's Base Pawn, regardless of your other pawns' positions
6
Special Pawn exclusive spaces
Some action spaces show a white cylinder icon and can only be used by Special Pawns
7
Voluntary action pass
Actions are never mandatory; you may voluntarily forgo performing the chosen action entirely
8
Abundance
Choose one Element from this section, remove it from the Action Area, and place it on any empty corner of any terrain tile on the map
9
Autotrophs
Either remove 1 Element from a Hydrothermal Vent tile that matches a type currently in this section, OR swap an Element in this section with one currently on a Hydrothermal Vent tile
10
Depletion
Choose any Element on the map that matches a type present in this section and return it to the bag
11
Adaptation
Choose 1 Element from this section and place it on an empty gray space on your Animal Board
12
Regression
Place one cube from your Gene Pool on an empty space to the left of the action space
13
Speciation
Choose any Element on the map matching the type shown on your chosen action space, then place cubes from your Gene Pool onto terrain tiles adjacent to that Element
14
Wanderlust (Exploration)
Choose a large terrain tile from one of three piles, flip the next tile face up, place the new tile on an empty hex adjacent to existing tiles
15
Tectonics
Choose a terrain tile on the grid's border, remove all species cubes temporarily, place a new Hydrothermal Vent tile on top (Submarine side in lower half, Land side in upper half)
16
Migration
Choose up to X of your species cubes anywhere on the map (where X matches the number shown on your action space) and move each to an adjacent terrain tile
17
Competition
Choose 1 tile on the map matching the terrain type shown on your action space that contains at least one of your species
18
Evolution
Choose exactly 1 tile on the map matching the terrain type shown, then score that tile
19
Domination
Choose exactly one of the six Element types your animal currently dominates, then take control of the associated Special Pawn
20
Thriving species
A species on a tile where at least one Element matches an Element on its Animal Board
21
Endangered species
A species on a tile where zero Elements match any Elements on its Animal Board
22
Extinction Event trigger
Occurs when a fish skeleton icon appears on a new Evolution card, and again immediately before final scoring
23
Survival Event trigger
Occurs when a fish icon appears on a new Evolution card, and again immediately before final scoring
24
Reseed Event trigger
Occurs when all player cubes occupy the right Food Chain space (except at end of game after Asteroid)

Winning

How to claim victory.

1
Game end trigger
When the Asteroid card is chosen during an Evolution action, the game will end at the conclusion of the current round
2
Complete the round
Continue play until the next Reseed Event would occur, then stop without performing the Reseed Event
3
Final Extinction Event
Perform one final Extinction Event, eliminating all endangered species
4
Final Survival Event
Perform one final Survival Event, awarding Bonus VP to the Survival card holder
5
Special Pawn VP bonus
Each player gains VP equal to the sum of Threshold marker positions for all Special Pawns they control
6
Final tile scoring
Score every terrain tile on the map one final time using standard Evolution scoring rules
7
Determine winner
The player with the highest total VP wins the game
8
Tiebreaker
If tied, the player whose animal is higher on the Food Chain wins (Reptiles, then Cephalopods, then Fish, then Crustaceans)

watch & Learn

Let the pros teach you with these top videos.

Dominant Species Marine - Présentation et Règles

Dominant Species Marine - Présentation et Règles

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優勢物種-海洋版(Dominant Species: Marine)

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FULL rules

Looking for the full details? Grab the complete rulebook as a downloadable PDF.

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